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This page explains in detail the interactions between the Mario series and the Metal Gear series.

Metal Gear Solid: The Twin Snakes

MGSTTS Mario Yoshi

Mario and Yoshi dolls in Metal Gear Solid: The Twin Snakes.

20040309 March 09, 2004

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A Mario and a Yoshi doll can be found in Otacon's lab. Shooting Mario makes the writing "1UP" appear while making the 1-Up sound. Shooting Yoshi causes him to say "Yoshi!"

MGSTTS Psycho Mantis-SMS

Psycho Mantis talking about Super Mario Sunshine.

Additionally, Psycho Mantis may mention Super Mario Sunshine during his battle if the player has a save file for that game in the memory card in slot 1.

License

Metal Gear Solid: The Twin Snakes was licensed by Nintendo to be released on the GameCube, so Konami was able to include references to Nintendo series, including Mario.